.container {
  width: min(calc(100% - 2rem), var(--container));
  margin-inline: auto;
}

.institution-bar {
  min-height: var(--institution-bar-height);
  display: flex;
  align-items: center;
  color: var(--color-on-strong-muted);
  background: var(--color-brand-950);
  font-size: var(--font-size-xs);
  line-height: 1.35;
}

.institution-bar__inner,
.institution-bar > .container {
  min-height: var(--institution-bar-height);
  display: flex;
  align-items: center;
  justify-content: space-between;
  gap: var(--space-3);
}

.institution-bar a {
  min-height: var(--target-size);
  display: inline-flex;
  align-items: center;
  color: var(--color-on-strong);
}

.institution-bar p {
  margin: 0;
}

.institution-bar__meta {
  display: none;
}

.site-header {
  position: sticky;
  z-index: var(--z-header);
  top: 0;
  min-height: var(--header-height);
  border-bottom: 1px solid var(--color-line);
  background: color-mix(in srgb, var(--color-surface) 96%, transparent);
  box-shadow: var(--shadow-xs);
  backdrop-filter: blur(12px);
}

.site-header__inner {
  min-height: var(--header-height);
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  align-items: center;
  gap: var(--space-2);
}

.site-header__actions {
  grid-column: 2;
  display: flex;
  align-items: center;
  gap: var(--space-2);
}

body.menu-open .site-header {
  z-index: var(--z-nav);
  -webkit-backdrop-filter: none;
  backdrop-filter: none;
}

body.menu-open .site-header__actions {
  position: relative;
  z-index: calc(var(--z-nav) + 1);
}

.brand {
  min-width: 0;
  display: inline-flex;
  align-items: center;
  gap: var(--space-3);
  color: var(--color-ink);
  text-decoration: none;
}

.brand:hover {
  color: var(--color-ink);
}

.brand__mark {
  width: 2.75rem;
  height: 2.75rem;
  flex: 0 0 2.75rem;
  object-fit: contain;
}

.brand .brand__mark--original {
  width: 4.75rem;
  height: auto;
  flex: 0 0 4.75rem;
  aspect-ratio: 251 / 141;
}

.brand__copy {
  min-width: 0;
  display: flex;
  flex-direction: column;
  line-height: 1.15;
}

.brand__copy strong {
  color: var(--color-link);
  font-size: 1.1rem;
  font-weight: 850;
  letter-spacing: 0.055em;
}

.brand__copy small {
  max-width: 21rem;
  overflow: hidden;
  color: var(--color-muted);
  font-size: 0.65rem;
  font-weight: 650;
  text-overflow: ellipsis;
  white-space: nowrap;
}

.brand__domain {
  display: none;
  color: var(--color-link);
  font-size: 0.625rem;
  font-weight: 850;
  letter-spacing: 0.11em;
}

.nav-toggle,
.theme-toggle {
  min-width: var(--target-size);
  min-height: var(--target-size);
  display: inline-grid;
  place-items: center;
  flex: 0 0 var(--target-size);
  padding: 0;
  border: 1px solid var(--color-line-strong);
  border-radius: var(--radius-sm);
  color: var(--color-link);
  background: var(--color-surface-raised);
  transition:
    color var(--duration-fast) var(--ease-standard),
    border-color var(--duration-fast) var(--ease-standard),
    background var(--duration-fast) var(--ease-standard);
}

.nav-toggle,
.theme-toggle {
  display: none;
}

.js .nav-toggle,
.js .theme-toggle {
  display: inline-grid;
}

.nav-toggle:hover,
.theme-toggle:hover {
  border-color: var(--color-link);
  color: var(--color-link);
  background: var(--color-brand-50);
}

.nav-toggle svg,
.theme-toggle svg,
.submenu-toggle svg {
  fill: none;
  stroke: currentColor;
  stroke-width: 1.8;
  stroke-linecap: round;
  stroke-linejoin: round;
}

.nav-toggle svg,
.theme-toggle svg {
  width: 1.25rem;
  height: 1.25rem;
}

.submenu-toggle {
  min-width: var(--target-size);
  min-height: var(--target-size);
  display: inline-grid;
  place-items: center;
  flex: 0 0 var(--target-size);
  padding: 0;
  border: 0;
  border-radius: var(--radius-sm);
  color: var(--color-muted);
  background: transparent;
  box-shadow: none;
  transition: color var(--duration-fast) var(--ease-standard);
}

.submenu-toggle:hover {
  color: var(--color-link);
  background: transparent;
}

.submenu-toggle svg {
  width: 0.875rem;
  height: 0.875rem;
}

.theme-toggle__moon,
.nav-toggle__close {
  display: none;
}

:root[data-theme='dark'] .theme-toggle__sun,
.nav-toggle[aria-expanded='true'] .nav-toggle__menu {
  display: none;
}

:root[data-theme='dark'] .theme-toggle__moon,
.nav-toggle[aria-expanded='true'] .nav-toggle__close {
  display: block;
}

.nav-backdrop {
  position: fixed;
  z-index: var(--z-backdrop);
  inset: 0;
  visibility: hidden;
  opacity: 0;
  border: 0;
  background: var(--color-overlay);
  transition: opacity var(--duration-normal) var(--ease-standard), visibility var(--duration-normal);
}

.nav-backdrop.is-open,
body.menu-open .nav-backdrop {
  visibility: visible;
  opacity: 1;
}

.primary-nav {
  grid-column: 1 / -1;
  position: static;
  visibility: visible;
  pointer-events: auto;
}

.primary-nav__panel {
  position: static;
  width: 100%;
  height: auto;
  padding: 0 0 var(--space-4);
  overflow: visible;
  border: 0;
  background: transparent;
  box-shadow: none;
  transform: none;
}

.js .primary-nav {
  position: fixed;
  z-index: var(--z-nav);
  inset: 0;
  grid-column: auto;
  grid-row: auto;
  width: 100%;
  height: 100vh;
  height: 100dvh;
  visibility: hidden;
  overflow: hidden;
  pointer-events: none;
}

.js .primary-nav__panel {
  position: absolute;
  inset: 0 0 0 auto;
  width: min(90vw, 22.5rem);
  height: 100vh;
  height: 100dvh;
  padding: max(calc(var(--institution-bar-height) + var(--header-height) + var(--space-4)), env(safe-area-inset-top)) var(--space-4) max(var(--space-5), env(safe-area-inset-bottom));
  overflow-y: auto;
  overscroll-behavior: contain;
  border-left: 1px solid var(--color-line);
  background: var(--color-surface);
  box-shadow: var(--shadow-md);
  transform: translateX(100%);
  transition: transform var(--duration-normal) var(--ease-standard);
}

.js .primary-nav.is-open {
  visibility: visible;
  overflow: visible;
  pointer-events: none;
}

.js .primary-nav.is-open .primary-nav__panel {
  transform: translateX(0);
  pointer-events: auto;
}

.primary-nav__list,
.submenu {
  margin: 0;
  padding: 0;
  list-style: none;
}

.primary-nav__list {
  display: grid;
  gap: var(--space-1);
}

.nav-item {
  min-width: 0;
  width: 100%;
}

.nav-item__row {
  display: flex;
  align-items: center;
  gap: 0;
  width: 100%;
}

.nav-link {
  min-height: var(--target-size);
  display: flex;
  align-items: center;
  flex: 1;
  padding: 0.65rem 0;
  color: var(--color-text);
  font-size: var(--font-size-sm);
  font-weight: 700;
  text-decoration: none;
}

.nav-link:hover,
.nav-link.is-active,
.nav-link[aria-current='page'],
.nav-item.is-active > .nav-item__row > .nav-link {
  color: var(--color-link);
}

.nav-link[aria-current='page'],
.nav-item.is-active > .nav-item__row > .nav-link {
  font-weight: 800;
}

.submenu-toggle svg {
  transition: transform var(--duration-normal) var(--ease-standard);
}

.submenu-toggle[aria-expanded='true'] svg {
  transform: rotate(180deg);
}

.submenu {
  display: grid;
  grid-template-columns: minmax(0, 1fr);
  gap: 0;
  width: 100%;
  margin: 0 0 var(--space-2);
  padding-left: 1.25rem;
  overflow: hidden;
}

.submenu:not([hidden]) {
  animation: submenu-reveal var(--duration-fast) var(--ease-standard) both;
}

.submenu a {
  min-height: var(--target-size);
  display: flex;
  align-items: center;
  padding: 0.45rem 0;
  border: 0;
  color: var(--color-muted);
  font-size: 0.8125rem;
  font-weight: 620;
  text-decoration: none;
}

.submenu a:hover,
.submenu a:focus-visible {
  color: var(--color-link);
  background: transparent;
}

@keyframes submenu-reveal {
  from {
    max-height: 0;
    opacity: 0;
  }

  to {
    max-height: 24rem;
    opacity: 1;
  }
}

.section {
  position: relative;
  padding-block: var(--space-8);
}

.section--soft {
  background: var(--color-surface-soft);
}

.section--ink {
  color: var(--color-on-strong-muted);
  background: var(--color-brand-950);
}

.section--ink :where(h1, h2, h3, h4, strong) {
  color: var(--color-on-strong);
}

.section--ink .eyebrow {
  color: var(--color-on-strong-accent);
}

.content-grid,
.card-grid,
.species-grid,
.webinar-grid,
.team-grid,
.contact-grid {
  display: grid;
  gap: var(--space-5);
}

.content-grid {
  align-items: start;
}

.section-heading {
  max-width: var(--measure);
  margin-bottom: var(--space-6);
}

.section-heading > * + * {
  margin-top: var(--space-3);
}

.site-footer {
  position: relative;
  padding: var(--space-8) 0 var(--space-5);
  color: var(--color-on-strong-muted);
  background: var(--color-brand-950);
}

.site-footer::before {
  position: absolute;
  inset: 0 0 auto;
  height: 3px;
  background: linear-gradient(90deg, var(--color-brand-500), var(--color-science-600));
  content: '';
}

.site-footer__grid {
  display: grid;
  gap: var(--space-6);
}

.footer-brand {
  max-width: 24rem;
}

.brand--footer {
  width: fit-content;
}

.footer-brand .brand__copy strong,
.site-footer strong,
.site-footer h2,
.site-footer h3 {
  color: var(--color-on-strong);
}

.footer-brand .brand__copy small {
  color: var(--color-on-strong-muted);
}

.footer-brand > * + * {
  margin-top: var(--space-4);
}

.footer-column {
  display: flex;
  flex-direction: column;
  align-items: flex-start;
  gap: var(--space-2);
}

.footer-column ul {
  display: grid;
  gap: var(--space-1);
  margin: 0;
  padding: 0;
  list-style: none;
}

.footer-column h2,
.footer-column h3 {
  margin-bottom: var(--space-2);
  font-size: var(--font-size-sm);
  letter-spacing: 0.08em;
  text-transform: uppercase;
}

.footer-column a {
  min-height: 2.25rem;
  display: inline-flex;
  align-items: center;
  color: var(--color-on-strong-muted);
  font-size: var(--font-size-sm);
  text-decoration: none;
}

.footer-column a:hover {
  color: var(--color-on-strong);
  text-decoration: underline;
}

.footer-bottom {
  display: grid;
  gap: var(--space-4);
  margin-top: var(--space-7);
  padding-top: var(--space-5);
  border-top: 1px solid rgba(255, 255, 255, 0.16);
  color: var(--color-on-strong-muted);
  font-size: var(--font-size-xs);
}

.footer-bottom p {
  margin: 0;
}

.footer-domain {
  color: var(--color-brand-300);
  font-size: var(--font-size-xs);
  font-weight: 850;
  letter-spacing: 0.12em;
}

.footer-build {
  color: var(--color-on-strong-muted);
  font-variant-numeric: tabular-nums;
}

.back-to-top {
  min-height: var(--target-size);
  display: inline-flex;
  align-items: center;
  justify-content: center;
  gap: var(--space-2);
  width: fit-content;
  padding: 0.6rem 0.85rem;
  border: 1px solid rgba(255, 255, 255, 0.28);
  border-radius: var(--radius-sm);
  color: var(--color-on-strong);
  background: rgba(255, 255, 255, 0.06);
  font-weight: 750;
  text-decoration: none;
}

.back-to-top:hover {
  color: var(--color-on-strong);
  background: rgba(255, 255, 255, 0.12);
}