@media (max-width: 23.99rem) {
  .container {
    width: min(calc(100% - 1.5rem), var(--container));
  }

  .brand {
    gap: var(--space-2);
  }

  .brand__mark {
    width: var(--target-size);
    height: var(--target-size);
    flex-basis: var(--target-size);
  }

  .brand__copy small {
    display: none;
  }

  .home-hero h1 {
    font-size: clamp(2rem, 9.5vw, 2.35rem);
  }

  .hero-actions .button,
  .page-actions .button {
    width: 100%;
  }

  .footer-bottom {
    align-items: start;
  }
}

@media (min-width: 40rem) {
  .container {
    width: min(calc(100% - 3rem), var(--container));
  }

  .institution-bar__meta {
    display: block;
  }

  .hero-visual {
    width: min(100%, 34rem);
  }

  .card-grid,
  .webinar-grid,
  .team-grid,
  .contact-grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .species-grid {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .cta-panel {
    grid-template-columns: minmax(0, 1fr) auto;
  }

  .site-footer__grid {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .footer-brand {
    grid-column: 1 / -1;
  }

  .footer-bottom {
    grid-template-columns: minmax(0, 1fr) auto;
    align-items: center;
  }

  .footer-bottom .back-to-top {
    justify-self: end;
  }

  .contact-form {
    grid-template-columns: repeat(2, minmax(0, 1fr));
  }

  .form-field--wide,
  .form-actions,
  .form-status {
    grid-column: 1 / -1;
  }
}

@media (min-width: 50rem) {
  .section {
    padding-block: var(--space-9);
  }

  .page-hero {
    padding: var(--space-7) 0 var(--space-9);
  }

  .page-hero__grid {
    grid-template-columns: repeat(auto-fit, minmax(min(100%, 24rem), 1fr));
    gap: var(--space-7);
  }

  .content-grid {
    grid-template-columns: minmax(0, 1.15fr) minmax(16rem, 0.85fr);
    gap: var(--space-7);
  }

  .card-grid--three {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .process-steps {
    grid-template-columns: repeat(auto-fit, minmax(min(100%, 15rem), 1fr));
  }

  .event-card {
    min-height: 19rem;
  }

  .event-card__media,
  .event-card > img {
    height: 100%;
    min-height: 100%;
    aspect-ratio: auto;
  }

  .team-card {
    grid-template-columns: auto minmax(0, 1fr);
    column-gap: var(--space-4);
  }

  .site-footer__grid {
    grid-template-columns: 1.4fr repeat(3, minmax(0, 1fr));
  }

  .footer-brand {
    grid-column: auto;
  }
}

@media (min-width: 64rem) {
  :root {
    --header-height: 4.5rem;
  }

  .site-header__inner {
    grid-template-columns: auto minmax(0, 1fr) auto;
    gap: var(--space-4);
  }

  .site-header__actions {
    grid-column: 3;
    grid-row: 1;
  }

  .brand__copy small {
    display: none;
  }

  .js .nav-toggle,
  .nav-backdrop {
    display: none;
  }

  .primary-nav,
  .primary-nav.is-open,
  .js .primary-nav,
  .js .primary-nav.is-open {
    position: static;
    z-index: auto;
    inset: auto;
    width: auto;
    height: auto;
    grid-column: 2;
    grid-row: 1;
    visibility: visible;
    overflow: visible;
    pointer-events: auto;
  }

  .primary-nav__panel,
  .js .primary-nav__panel {
    position: static;
    width: auto;
    height: auto;
    padding: 0;
    overflow: visible;
    border: 0;
    background: transparent;
    box-shadow: none;
    transform: none;
    transition: none;
  }

  .primary-nav__list {
    display: flex;
    align-items: center;
    justify-content: flex-end;
    gap: 0.125rem;
  }

  .nav-item {
    position: relative;
    width: auto;
  }

  .nav-item__row {
    width: auto;
  }

  .nav-link {
    position: relative;
    min-width: 2.875rem;
    min-height: 2.75rem;
    justify-content: center;
    flex: 0 1 auto;
    padding-inline: 0.55rem;
    font-size: 0.78rem;
    white-space: nowrap;
  }

  .nav-item--group .nav-link {
    min-width: var(--target-size);
    padding-right: 0.15rem;
  }

  .nav-link[aria-current='page'],
  .nav-item.is-active > .nav-item__row > .nav-link {
    box-shadow: none;
  }

  .nav-link[aria-current='page']::after,
  .nav-item.is-active > .nav-item__row > .nav-link::after {
    content: '';
    position: absolute;
    right: 0.5rem;
    bottom: 0.2rem;
    left: 0.5rem;
    height: 2px;
    border-radius: 999px;
    background: currentColor;
  }

  .submenu-toggle {
    min-width: var(--target-size);
    min-height: var(--target-size);
    flex-basis: var(--target-size);
    place-items: center start;
    border: 0;
    background: transparent;
    box-shadow: none;
  }

  .submenu {
    position: absolute;
    z-index: 10;
    inset: calc(100% + 0.4rem) auto auto 0;
    width: clamp(16.25rem, 20vw, 20rem);
    margin: 0;
    padding: 0.45rem;
    overflow: hidden;
    border: 1px solid var(--color-line);
    border-radius: var(--radius-md);
    background: var(--color-surface-raised);
    box-shadow: var(--shadow-sm);
  }

  .submenu a {
    padding: 0.6rem 0.7rem;
    border: 0;
    border-radius: var(--radius-sm);
  }

  .home-hero {
    padding: var(--space-8) 0 var(--space-9);
  }

  .home-hero__grid,
  .home-hero > .container {
    grid-template-columns: minmax(0, 1.08fr) minmax(22rem, 0.92fr);
    gap: clamp(3rem, 6vw, 5rem);
  }

  .hero-visual {
    justify-self: end;
  }

  .card-grid,
  .webinar-grid {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .card-grid--four {
    grid-template-columns: repeat(4, minmax(0, 1fr));
  }

  .card-grid:has(> .event-card:only-child),
  .webinar-grid:has(> .webinar-card:only-child) {
    grid-template-columns: minmax(0, 52rem);
  }

  .team-grid {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .contact-grid {
    grid-template-columns: repeat(3, minmax(0, 1fr));
  }

  .site-footer__grid {
    grid-template-columns: minmax(14rem, 1.55fr) repeat(4, minmax(0, 1fr));
    gap: var(--space-6);
  }
}

@media (min-width: 72rem) {
  .brand__copy small,
  .brand__domain {
    display: block;
  }

  .brand__domain {
    margin-top: 0.18rem;
  }

  .nav-item--direct .nav-link {
    padding-inline: 0.62rem;
  }

  .nav-item--group .nav-link {
    padding-left: 0.62rem;
    padding-right: 0.15rem;
  }

  .nav-link {
    font-size: var(--font-size-sm);
  }
}

@media (min-width: 85rem) {
  .home-hero__grid,
  .home-hero > .container {
    gap: 5.5rem;
  }
}

@media (max-height: 36rem) and (max-width: 63.99rem) {
  .primary-nav__panel {
    padding-top: calc(var(--institution-bar-height) + var(--header-height) + var(--space-2));
    padding-bottom: var(--space-3);
  }

  .nav-link,
  .submenu a {
    min-height: var(--target-size);
    padding-block: 0.45rem;
  }
}

@media (forced-colors: active) {
  .button,
  .nav-toggle,
  .theme-toggle,
  .submenu-toggle,
  .card,
  .project-card,
  .resource-card,
  .event-card,
  .webinar-card,
  .team-card,
  .contact-card {
    border: 1px solid CanvasText;
  }

  .nav-link[aria-current='page'],
  .nav-item.is-active > .nav-item__row > .nav-link {
    outline: 2px solid Highlight;
    outline-offset: -2px;
  }
}